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SOunds like a driver problem then or something ingame has set the sound volume to 0 during gameplay. Maybe another software or sound related software you got installed. Try to alt+tab back to desktop and check if the sound slider for ESO is set to maximum and if the active sound harware is your soundcard + boxes / headset, and not any monitor (which got no boxes attached currently).
Sometimes nvidia drivers e.g. install HDMI audio drivers and the the active sound card is the monitor then all of sudden instead of your boxes/headset.
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